Nonequilibrium models of optimal enhancer function
R. Grah, B. Zoller, G. Tkadik, PNAS 117 (2020) 31614–31622.

Gene amplification as a form of population-level gene expression regulation
I. Tomanek, R. Grah, M. Lagator, A.M.C. Andersson, J.P. Bollback, G. Tkadik, C.C. Guet, Nature Ecology &; Evolution 4 (2020) 612–625.

Normative models of enhancer function
R. Grah, B. Zoller, G. Tkadik, BioRxiv (2020).

Gene regulation across scales – how biophysical constraints shape evolution
R. Grah, Gene Regulation across Scales – How Biophysical Constraints Shape Evolution, Institute of Science and Technology Austria, 2020.