Gene regulation across scales – how biophysical constraints shape evolution
R. Grah, Gene Regulation across Scales – How Biophysical Constraints Shape Evolution, Institute of Science and Technology Austria, 2020.

Nonequilibrium models of optimal enhancer function
R. Grah, B. Zoller, G. Tkadik, PNAS 117 (2020) 31614–31622.

Gene amplification as a form of population-level gene expression regulation
I. Tomanek, R. Grah, M. Lagator, A.M.C. Andersson, J.P. Bollback, G. Tkadik, C.C. Guet, Nature Ecology &; Evolution 4 (2020) 612–625.

Normative models of enhancer function
R. Grah, B. Zoller, G. Tkadik, BioRxiv (2020).